Crawley joined an exclusive band of non-league clubs to reach the fifth round of the FA Cup with a 1-0 win over League Two Torquay at Plainmoor.
Matt Tubbs fired the visitors ahead six minutes before the break, and after the Gulls had Chris Zebroski sent off for a second bookable offence, Tubbs and team-mate Craig McAllister had second-half penalties saved by Scott Bevan.
Crawley had defender Dean Howell sent off in the 83rd minute and almost paid the price late on when defender Kevin Nicholson crashed a shot against the crossbar, but they managed to hang on to claim a place in the last 16.
Swansea defender Alan Tate put through his own net in the 88th minute to hand Leyton Orient a 2-1 win and send them into the fifth round for the first time in 29 years.
Jimmy Smith headed Russell Slade's men ahead in the 35th minute, but the Swans levelled on the stroke of half-time through Cedric Van der Gun. Yet the real drama was still to come as Tate diverted a cross from substitute Paul-Jose M'Poku into his own net to send the League One club into the next round.
There was something of a shock at Vicarage Road too, where Watford of the Championship went down to Brighton of League One after keeper Rene Gilmartin made a mess of dealing with Ashley Barnes' 16th-minute shot.
League Two Stevenage, conquerors of Premier League Newcastle in the third round, were unable to extend their run further despite coming from behind at home to Reading.
Mikele Leigertwood put the visitors ahead with 23 minutes gone, but Darius Charles equalised with 18 minutes left, only for Shane Long to snatch victory for the Championship club at the death.
Burton's adventure also drew to a close at Burnley as a Chris Eagles double proved enough for the Clarets despite loan signing Calvin Zola's 80th-minute strike.
League Two Hereford took a ninth-minute lead against Sheffield Wednesday through Stuart Fleetwood, but Darren Potter's equaliser six minutes later set the scene for Clinton Morrison to score twice from the penalty spot, either side of Jermaine Johnson's 77th-minute strike as the Owls ran out 4-1 winners.
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