Jonny Gray and Ruaridh Jackson return to a Scotland side showing five changes from the end of the Six Nations for Saturday's first Test against Japan.
All but one of the alterations are injury-related, with four players unavailable and Gray returning to partner brother Richie at lock after missing the match against Ireland in Dublin.
The only unenforced change had been highlighted in advance with Duncan Weir, who started in Ireland, failing to make the tour, meaning a return for Jackson at fly-half a few days after he confirmed his move from Wasps to Harlequins.
There is a surprise selection on the wing, where coach Vern Cotter opted to replace the injured Tim Visser with Edinburgh's Damien Hoyland - a reward for a profitable club season where he finished second in the Guinness PRO12 try-scoring table - rather than Saracens-bound Sean Maitland.
The other changes see Matt Scott come in for Alex Dunbar at centre while Ross Ford has a tight calf muscle and drops out with Stuart McInally coming in as his replacement at hooker.